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WASHINGTON — Attorney General Eric Holder on Tuesday expressed regret to the family of a slain federal border agent as the attorney general underwent pointed questioning by Senate Republicans about a flawed arms trafficking investigation called Operation Fast and Furious.

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Holder said that he can only imagine the pain that the family of border agent Brian Terry feels. The attorney general added, however, that it is not fair to assume that mistakes in Operation Fast and Furious led to Terry’s death.

Two of the more than 2,000 guns involved in Operation Fast and Furious turned up at the scene of a shootout in Arizona that resulted in the death of Terry, a Customs and Border Protection agent.